The apoplast is the non-living part of plant tissue that enables movement of water and solutes through cell walls and intercellular spaces without crossing cell membranes. It plays a key role in water transport, nutrient movement, and transpiration — an essential concept often compared with the symplast pathway.
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The apoplast is the continuous system of cell walls and intercellular spaces in plant tissues that allows for the movement of water and solutes outside of the cell membranes. It forms a crucial pathway for water and nutrient transport in plants because it provides a low-resistance pathway for water movement from roots to leaves. It allows for rapid, long-distance water transport without crossing cell membranes, which is essential for maintaining plant hydration and supporting transpiration.
The major components of apoplast include:
Components | Functions |
Provide porous structure for water movement | |
Intercellular spaces | Facilitate diffusion between adjacent cells |
Xylem vessels | Enable long distance transport of water and minerals |
Plants require water to survive just as animals and mammals need. Plants require water to complete their daily basis of biological processes. Water plays a very important role in processes such as seed germination and photosynthesis. Three major water means of transport are discussed below:
Apoplast is defined as the pathway in which the movement of water occurs through the cell wall and other intercellular spaces that are present in the cell. The apoplast, also known as the cell wall of the plant, is located on the outer side of the cell.
Symplast is defined as the transfer of water from one cell to another cell with the help of some connections that are called plasmodesmata. The inner side of the plasma membrane consists of symplast.
As the name suggests, transmembrane is defined as a combination of both the Apoplast and Symplast pathways and forms a transmembrane pathway. This is the pathway that helps in the movement of water across cells and cell walls.
Materials required during the Apoplast Pathway are discussed below in the table:
Materials Needed | Explanation |
Oxygen |
|
Carbon dioxide |
|
Organic Nutrients |
|
Inorganic ions and water |
|
The functions of apoplast in plants include:
It enables the rapid movement of water and solute across the cell walls.
It also maintains the turgor and hydration in the plant tissues.
It facilitates the process of guttation and transpiration in plants.
It provides mechanical support because of the presence of lignified xylem.
The differences between the apoplast and symplast pathway are:
Apoplast Pathway | Symplast Pathway |
Apoplast pathway is the movement of water through adjacent cell walls. | Symplast pathway, water moves from one cell to another through plasmodesmata. |
Movement through apoplast does not involve crossing the cell membrane. | During symplast movement, the water travels through cells. |
Movement is faster. | It is relatively slower. |
It does not provide any barrier in water movement. | Water molecules are unable to penetrate casparian strips. |
It is not aided by the streaming movement of cytoplasm. | It is aided by the streaming movement of cytoplasm. |
The metabolic rate of the cells in the root cortex does not affect the movement of water. | The metabolic rate of the cells in the root cortex highly affects the movement of water. |
The basic transportation of water and ions through the apoplast path includes:
Xylem vessels and tracheids act as continuous channels.
The lignin depositions prevent collapse under tension.
It also helps to maintain an uninterrupted water column during transpiration.
The key concepts to be covered under this topic for different exams are:
Mechanism of Root pressure
Factors affecting Root pressure
Q1. The apoplast pathway involves movement of water through:
Cytoplasm
Vacuoles
Cell walls and intercellular spaces
Plasmodesmata
Correct answer: 3) Cell walls and intercellular spaces
Explanation:
The apoplast pathway refers to the movement of water through the cell walls and intercellular spaces without crossing the plasma membrane. It is a non-living continuum outside the protoplast. Since it does not involve the cytoplasm, vacuoles, or plasmodesmata (which are cytoplasmic connections used in the symplast pathway), the correct answer is cell walls and intercellular spaces. This makes apoplast transport rapid and passive.
Hence, the correct answer is option 3) Cell walls and intercellular spaces.
Q2. Which of the following is absent in apoplast transport?
Cell membrane crossing
Passive movement
Intercellular spaces
Diffusion
Correct answer: 1) Cell membrane crossing
Explanation:
In apoplast transport, water and solutes move outside the cell membranes through cell walls and intercellular spaces, so they do not cross the plasma membrane until they reach the endodermis (Casparian strip). Passive movement, intercellular spaces, and diffusion are all part of apoplast transport. Therefore, the correct answer is cell membrane crossing, which is absent until the symplast pathway takes over at the endodermis.
Hence, the correct answer is option 1) Cell membrane crossing.
Q3. Lignin in xylem vessels mainly provides:
Photosynthesis
Structural support and water transport
Respiration
Nutrient storage
Correct answer: 2) Structural support and water transport
Explanation:
Lignin is a complex organic polymer deposited in the cell walls of xylem vessels. It provides mechanical strength to withstand the negative pressure generated during transpiration pull. Additionally, lignin makes the walls impermeable to water leakage, creating an efficient, continuous channel for long-distance water transport. It does not play a role in photosynthesis, respiration, or direct nutrient storage, making structural support and water transport the correct answer.
Hence, the correct answer is option 2) Structural support and water transport.
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Frequently Asked Questions (FAQs)
The term apoplast was coined by a german plant physiologist named Munch.
The main significance of apoplast pathway are :
It helps in the interaction of plant with the outer environment
It helps to provide resistance to plant against some toxic substances
It helps in eliminating aluminum ions which cause toxicity.
The cells of the plant require water and many other inorganic ions for the proper growth and these are transported from the soil with the help of the root. The roots follow the apoplastic pathway for the transport of water and ions.